Published & in Press

*student co-author,

mentee research project

(1) Olsen KC, Ryan WH, Kosman ET, Moscoso JA, Levitan DR, Winn AA. 2021. Lessons from the study of plant mating systems for exploring the causes and consequences of inbreeding in marine invertebrates. Marine Biology, 168: article 39. (PDF)

(2) Cahill R*, Krueger-Hadfield SA, Ryan WH. 2021. The effects of stress on the bacterial community associated with the sea anemone Diadumene lineata. Journal of Emerging Investigators, 3: 1-7. (PDF)

(3) Ryan WH, Aida J*, Krueger-Hadfield SA. 2021. The contribution of clonality to population genetic structure in the sea anemone Diadumene lineata. Journal of Heredity, 112: 122 - 139. (PDF) 

(4) Krueger-Hadfield SA and Ryan WH. Influence of exogenous nutrients on ploidy-specific performance in an invasive, haplodiplontic red macroalga. 2020. Journal of Phycology, 56(4): 1114 - 1120. (PDF) https://doi.org/10.1111/jpy.13011

(5) Olsen K, Ryan WH, Winn AA, Kosman ET, Moscoso JA, Krueger-Hadfield SA, Burgess SC, Carlon D, Grosberg RK, Kalisz S, and Levitan DR. 2020. Inbreeding shapes the evolution of marine invertebrate mating systems. Evolution 74(5): 871 - 882 (PDF)

(6) Bedgood SA, Bracken MES, Ryan WH, Levell ST , Wulff JL. 2020. Environmental drivers of adult locomotion and reproduction in a symbiont-hosting sea anemone. Marine Biology 161:39 (PDF)

(7) Ryan WH, L Adams, G Bonthond, N Mieszkowska, KE Pack, and SA Krueger-Hadfield. 2019. Environmental regulation of individual body size contributes to geographic variation in clonal life cycle expression. Marine Biology 166: 157 - 173 (PDF)

(8) Ryan WH, S Heiser*, MD Curtis*, CD Amsler, T Bayer, G Bonthond*, G Wang, F Weinberger, and SA Krueger-Hadfield. 2019. A method for using photographic color information as a proxy for pigment composition in macroalgaeCryptogamie Algologie 40 (7): 73 - 83 (PDF)

(9) Ryan WH and TE Miller. 2019. Geographic patterns of size, density, and gametogenesis in a clonal sea anemone across the Atlantic coast of North America. Marine Ecological Progress Series 611: 129-141 (Email for PDF)   https://doi.org/10.3354/meps12862

(10) Krueger-Hadfield SA, Stephens TA, Ryan WH, and Heiser S. 2018. Everywhere you look, everywhere you go, there’s an estuary invaded by the red seaweed Gracilaria vermiculophylla. Bioinvasion Records. 7 (4): 343 – 355 (PDF)

(11) Ryan WH. 2018. Seasonal patterns and geographic variation in temperature dependent fission rate plasticity in a clonal sea anemone. The American Naturalist 192: 210 - 219 (PDF(Appendix)

(12) Burgess S, Ryan W, Blackstone N, Edmunds P, Hoogenboom M, Levitan D and Wulff J. 2017. What can modular marine invertebrates tell us about variation in metabolic scaling? Invertebrate Biology 136: 456 - 472 (PDF)

(13) Ryan WH, Gornish ES, Christenson L, Halpern S, Henderson S, Lebuhn G and Miller TE. 2017. Initiating and managing long-term data with amateure scientists. The American Biology Teacher 79(1): 30-36 (PDF)

(14) Ryan WH and Kubota S. 2016. Morphotype distribution of the sea anemone Diadumene lineata in Tanabe Bay, Wakayama: a comparison with Uchida (1936) after 80 years. Publications of the Seto Marine Biological Laboratory 44: 1-6 (PDF)

(15) Pastore AI, Prather CM, Gornish ES, Ryan WH, Ellis RD and Miller, TE. 2014. Testing the competition-colonization trade-off with a 32 year study of saxicolous lichen community.  Ecology 95:  306-15 (PDF)

Non-peer reviewed and informal science writing

Ryan WH. Apr 2021. Behind the science: looking for the population genetic signatures of variable clonality across an environmental gradient. American Genetic Association Blog. (Link)

Ryan WH. May 2020. Integrating patterns of growth, sex, and clonal reproduction over time and space. RCN for Evolution in Changing Seas Blog. (Link)

Scholl JP, WH Ryan and J Mutz. 2016. Animating the primary literature for students and other curious people. The Bulletin of the Ecological Society of America 97(2): 192-199 (PDF)